For example, you can take those exit pages a little deeper and go through all of your content to see if anything needs an update or rewrite. While you’re doing this, you should also see if you can combine a few posts into single “mega” posts that are more likely to rank higher than each of the smaller posts, coming up with a solid content marketing plan will help you plan content for the future which will help increase your organic traffic further.
Google claims their users click (organic) search results more often than ads, essentially rebutting the research cited above. A 2012 Google study found that 81% of ad impressions and 66% of ad clicks happen when there is no associated organic search result on the first page. Research has shown that searchers may have a bias against ads, unless the ads are relevant to the searcher's need or intent 
Like you I am a scientist and like you did in the past, I am currently working on translating great scientific literature into tips. In my case it’s child development research into play tips for parents. I can already see that the outcome of my experiment is going to be the same as yours. Great content but who cares. I hadn’t even thought about my key influences. I know some important ones, but don’t see how they would share my content. I thought I was writing content for my potential customers. Is your SEO that works course the same as the content that gets results course? Sorry if I sound a bit dim asking that question.
But optimized keywords aren't just for organic search marketing. There's tremendous value in optimizing your PPC keywords. To be effective with your pay-per-click campaigns, you must speak the searcher's language. The more keywords you select that your customers are searching for, the more traffic you'll drive to your website, the more consumers you'll convert to customers.
“In conclusion, this research illuminates how content characteristics shape whether it becomes viral. When attempting to generate word of mouth, marketers often try targeting “influentials,” or opinion leaders (i.e., some small set of special people who, whether through having more social ties or being more persuasive, theoretically have more influence than others). Although this approach is pervasive,recent research has cast doubt on its value (Bakshy et al. 2011; Watts 2007) and suggests that it is far from cost effective. Rather than targeting “special” people, the current research suggests that it may be more beneficial to focus on crafting contagious content. By considering how psychological processes shape social transmission, it is possible to gain deeper insight into collective outcomes, such as what becomes viral.”
Same principles that goes to blog post commenting, also goes to forum marketing. Very simple - just try to genuinely get into discussions, share your opinion and help people. The great thing about forums is signature. You can add link to your website that will be shown below each of your replies and threads. This is another way to directly drive natural traffic to your website and build up backlinks for SEO at the same time. Look for forums that may have participants who could be interested in your online business. Niche forums are the best choice. Good example is Warrior Forum for internet marketers.
Plan your link structure. Start with the main navigation and decide how to best connect pages both physically (URL structure) and virtually (internal links) to clearly establish your content themes. Try to include at least 3-5 quality subpages under each core silo landing page. Link internally between the subpages. Link each subpage back up to the main silo landing page.